Graphoderus zonatus (Hoppe, 1795)
(Spangled Diving Beetle)

Taxonomic hierarchy:

Species	Graphoderus zonatus (Spangled Diving Beetle)
Genus	Graphoderus (a genus of diving beetles)
Subfamily	DYTISCINAE (a subfamily of water beetles)
Family	DYTISCIDAE (diving beetles)
Suborder	ADEPHAGA (water and ground beetles)
Order	COLEOPTERA (beetles)
Division	Endopterygota (bees, beetles, flies, moths and other insects with wings developing internally)
Infraclass	Neoptera (bees, beetles, flies, grasshoppers, moths and other advanced insects)
Subclass	PTERYGOTA (bees, beetles, dragonflies, flies, grasshoppers, moths and other winged insects)
Class	INSECTA (true insects)
Subphylum	HEXAPODA (insects and other 6-legged organisms)
Phylum	ARTHROPODA (arthropods)
Superphylum	ECDYSOZOA (skin shedders)
Clade	Bilateria (bilaterally symmetrical animals)
Subkingdom	EUMETAZOA (metazoans)
Kingdom	ANIMALIA (animals)
Domain	Eukaryota (eukaryotes)
Life	BIOTA (living things)

Graphoderus zonatus is also a member of:
water beetles - water beetles (via Dytiscidae)
Invertebrates - Invertebrates (via Arthropoda)

Identification Works

	Author	Year	Title	Source
	Cooter, J. & Barclay M.V.L. (eds)	2006	A Coleopterist’s handbook	The Amateur Entomolgist, 11, Fourth Edition edition, 439pp, Amateur Entomologists’ Society
	Woolmer Conservation Group	2006	Woolmer Forest: 30 years of conservation work	British Wildlife Vol 17, No. 3: 185-193.

Graphoderus zonatus (Spangled Diving Beetle) may also be included in identification literature listed under the following higher taxa:

DYTISCIDAE
(diving beetles)

ADEPHAGA
(water and ground beetles)

COLEOPTERA
(beetles)

INSECTA
(true insects)

HEXAPODA
(insects and other 6-legged organisms)

ARTHROPODA
(arthropods)

ANIMALIA
(animals)

Eukaryota
(eukaryotes)

BIOTA
(living things)

water beetles
(water beetles)

NBN (data.nbn.org.uk) has a distribution map for Graphoderus zonatus (Spangled Diving Beetle)

BioInfo (www.bioinfo.org.uk) has 0 host/parasite/foodplant and/or other relationships for Graphoderus zonatus (Spangled Diving Beetle)

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Taxonomic hierarchy:

Graphoderus zonatus is also a member of:

Identification Works

BioImages - Virtual Field-Guide (UK)